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anas, like other male examples of Squamata, have two hemipenes. During copulation, one hemipenis is inserted into the female's cloacal vent. A female can store sperm from previous mates for several years to continue to fertilize her eggs in case she finds no male within her territory when she is ready to lay again. Mating/courtship Iguanas tend to follow a promiscuous or polygynandrous mating style during the dry season. Mating during the dry season ensures that their offspring will hatch during the wet or rainy season when food will be more plentiful. Females control large territories, where they make several nests. Males compete for the females in an area and mark their won territory with a pheromone secreted from the femoral pores on the dorsal side of their hind limbs. Male behavior during sexual competition involves head bobbing, extending and retracting their dewlap, nuzzling and biting the necks of females, and on occasion, changing color. Once a female chooses a male, he straddles the female and holds her in place by biting onto her shoulder, which sometimes leaves scars on females. After copulation, eggs are laid within several nests and allowed to incubate. This low level of parental intervention with their offspring makes iguanas an example of r-strategy reproduction.[citation needed] Phylogeny A phylogeny based on nuclear protein-coding genes, reviewed by Vidal and Hedges (2009), suggested that the subclade Iguania is in a group with snakes and anguimorphs (lizards). These

 

 

 

 

 

 

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only phosphates and nitrates, although this varies with location. Prior to their being phasing out in the 1970's, phosphate-containing detergents contributed to eutrophication. Since then, sewage and agriculture have emerged as the dominant phosphate sources. The main sources of nitrogen pollution are from agricultural runoff containing fertilizers and animal wastes, from sewage, and from atmospheric deposition of nitrogen originating from combustion or animal waste. The limitation of productivity in any aquatic system varies with the rate of supply (from external sources) and removal (flushing out) of nutrients from the body of water. This means that some nutrients are more prevalent in certain areas than others and different ecosystems and environments have different limiting factors. Phosphorus is the limiting factor for plant growth in most freshwater ecosystems, and because phosphate adheres tightly to soil particles and sinks in areas such as wetlands and lakes, due to its prevalence nowadays more and more phosphorus is accumulating inside freshwater bodies. In marine ecosystems, nitrogen is the primary limiting nutrient; nitrous oxide (created by the combustion of fossil fuels) and its depos

 

 

 

 

 

 

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dae, when compared to habitat, is highly variable, although most every land habitat in the Americas has at least some of these birds. The habitats of tropical lowland evergreen forest and montane evergreen forest have the highest single site species diversity while many habitats including rivers, palm forest, white sand forest, tropical deciduous forest edge, southern temperate forest, southern temperate forest edge, semi-humid/humid montane scrub, and northern temperate grassland have the lowest single species diversity. The variation between the highest and the lowest is extreme; ninety species can be found in the tropical lowland evergreen forests while the number of species that can be found in the habitats listed above typically are in the single digits. This may be due in part to the fewer niches found in certain areas and therefore fewer places for the species to occupy. Tyrannidae specialization among habitats is very strong in tropical lowland evergreen forests and montane evergreen forests. These habitat types, therefore, display the greatest specialization. The counts differ by three species (tropical lowland evergreen forests have 49 endemic species and montane evergreen forests have 46 endemic species). It can be assumed that they both have similar levels of specialization. Regionally, the Atlantic Forest has the highest species richness with the Chocó follo

 

 

 

 

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dae, when compared to habitat, is highly variable, although most every land habitat in the Americas has at least some of these birds. The habitats of tropical lowland evergreen forest and montane evergreen forest have the highest single site species diversity while many habitats including rivers, palm forest, white sand forest, tropical deciduous forest edge, southern temperate forest, southern temperate forest edge, semi-humid/humid montane scrub, and northern temperate grassland have the lowest single species diversity. The variation between the highest and the lowest is extreme; ninety species can be found in the tropical lowland evergreen forests while the number of species that can be found in the habitats listed above typically are in the single digits. This may be due in part to the fewer niches found in certain areas and therefore fewer places for the species to occupy. Tyrannidae specialization among habitats is very strong in tropical lowland evergreen forests and montane evergreen forests. These habitat types, therefore, display the greatest specialization. The counts differ by three species (tropical lowland evergreen forests have 49 endemic species and montane evergreen forests have 46 endemic species). It can be assumed that they both have similar levels of specialization. Regionally, the Atlantic Forest has the highest species richness with the Chocó follo

 

 

 

 

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derwings' outer margins, corresponding to where the black border ends on the upperside. Females differ from the males in having yellow spots along the black borders on the upperside. In flight, Colias croceus is easily identifiable by the intense yellow colouring, much brighter than that of the lemon-yellow male common brimstone which also lacks black markings. Like all Colias species they never open their wings at rest. In about 5% of females, the golden upperside colouration is replaced by a pale cream colour. These females have been distinguished as form helice. The pale form helice does not seem to be distinct, as intermediates exist, and the variation is to some extent related to humidity during development, with dryer conditions producing paler colouration. These pale forms helice can be confused with Berger's clouded yellow (Colias alfacariensis) and the rarer pale clouded yellow (Colias hyale). Even the palest C. croceus tends to have more black on the upperside, however, in particular on the hindwings. Young caterpillars are yellow-green with a black head. Later they become completely dark green, with a white red-spotted lateral line after the third moult. The pupae are green and have a yellow side stripe. This species is similar to Colias myrmidone, Colias chrysotheme, Colias erate, Colias hyale, Colias alfacariensis, Colias caucasica, Colias aurorina. Life cycle and larval host plants Adults fly from March to October. In southern Europe and North Africa they breed continuously throughout the year. Eggs are laid singly on food plant leaves. Usually an extraordinary number of eggs – up to 600 – are laid from a single female. The caterpillars grow fast in warm weather, sometimes pupating within a mo

 

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directs here. For the comic book grant foundation, see Xeric Foundation. Desertic region at the M'Sila Province, Algeria Extent of deserts and xeric shrublands Deserts and xeric shrublands are a biome defined by the World Wide Fund for Nature. Deserts and xeric (Ancient Greek ξηρ?ς x?rós 'dry') shrublands form the largest terrestrial biome, covering 19% of Earth's land surface area. Ecoregions in this habitat type vary greatly in the amount of annual rainfall they receive, usually less than 250 millimetres (10 in) annually except in the margins. Generally evaporation exceeds rainfall in these ecoregions. Temperature variability is also diverse in these lands. Many deserts, such as the Sahara, are hot year-round, but others, such as East Asia's Gobi Desert, become quite cold during the winter. Temperature extremes are a characteristic of most deserts. High daytime temperatures give way to cold nights because there is no insulation provided by humidity and cloud cover. The diversity of climatic conditions, though quite harsh, supports a rich array of habitats. Many of these habitats are ephemeral in nature, reflecting the paucity and seasonality of available water. Woody-stemmed shrubs and plants characterize vegetation in these regions. Above all, these plants have evolved to minimize water loss. Animal biodiversity is equally well adapted and quite dive